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Table 1 Morphometric assessments of expanded leaves of winter and summer (±SD, n = 36)

From: Traits of seasonal dimorphism associated with adaptation to cold stress in Origanum dictamnus L. (Lamiaceae)

  Winter leaves Summer leaves
Leaf length (mm) 10.8 ± 3.1a 23.4 ± 8.8b
Leaf width (mm) 10.1 ± 2.5a 21.7 ± 6.9b
Leaf surface area (on each side) (mm2) 93.7 ± 6.7a 408.6 ± 49.2b
Leaf thickness (μm) 254.5 ± 25.6a 190.1 ± 17.9b
Relative volume percentage of leaf upper epidermis (leaf cross section) 12.8 ± 2.1a 12.5 ± 1.4a
Relative volume percentage of leaf palisade parenchyma (leaf cross section) 53.2 ± 9.3a 52.9 ± 10.6a
Relative volume percentage of spongy parenchyma (leaf cross section) 29.3 ± 4.7a 23.3 ± 1.8b
Relative volume percentage of leaf lower epidermis (leaf cross section) 5.8 ± 0.6a 4.1 ± 0.7b
Density of stomata on the upper leaf side (No mm−2) * *
Density of stomata on the lower leaf side (No mm−2) 564.4 ± 81.1a 458.2 ± 66.3b
Density of peltate glandular hairs on the upper leaf side (No mm−2) 3.5 ± 2.0a 4.6 ± 1.4b
Density of peltate glandular hairs on the lower leaf side (No mm−2) 4.8 ± 1.4a 6.1 ± 1.4b
  1. Means between columns with different letters are significantly different at p < 0.05.
  2. *Stomata on the upper leaf side are extremely limited.